, 2013). A lasting implication of this approach, which is sometimes also referred to as continuous cover irregular shelterwood (Raymond et al., 2009), is a substantial portion of the forest canopy is maintained across the stand throughout multiple rotations. Other

silvicultural approaches such as shelterwood Pictilisib cuts, where standing trees are left following harvest primarily to establish and promote regeneration, may however provide some de facto benefits for biodiversity at least in the short-term. In shelterwood silvicultural systems, standing trees are left for several years to maintain an abundant seed source and ensure successful regeneration following harvest at which point seed trees may then be harvested TSA HDAC chemical structure ( Lieffers et al., 2003). As well as potentially

leaving trees only temporarily (e.g. 10–20 years), shelterwood systems tend to leave as few trees as possible to ensure both adequate seed and light for regeneration as well as reduced risk of loss to windthrow ( Smith et al., 1997 and Nyland, 2002). As such, standard shelterwood systems often have lower levels of retention than multi-cohort approaches. In boreal systems, where limited topographic variation permits extensive access by harvesting machinery, dispersed retention targets (i.e. leave trees) within either shelterwood or multicohort management are often achieved through a series of residual vegetation strips, which may be thinned depending on prescription targets, and harvested corridors (David et al., 2000). If retention within residual vegetation strips is held relatively constant, clearly lower overall retention levels within a harvest unit will necessitate either wider or more harvested corridors. Larger harvested corridors may create significant edge effects into residual vegetation strips affecting microclimate (Zheng and Chen, 2000) and may harbor different species assemblages of plants (Craig and Macdonald, 2009),

fungi (Lazaruk et al., 2005) and animals (Lindo and Visser, 2004). Conversely, more open habitats, such as machine corridors, may serve as favorable Depsipeptide supplier habitats for generalist or disturbance-adapted species (Klimaszewski et al., 2005, Niemelä et al., 2007 and Brais et al., 2013). For organisms that respond to stand-level changes caused by forest management such as ground beetles (Coleoptera: Carabidae), post-harvest retention levels affect both overall abundance and species composition (Koivula, 2002, Martikainen et al., 2006, Halaj et al., 2008 and Work et al., 2008). Ground beetles are generalist predators which reside in forest leaf litter and have been widely used to evaluate the impacts of forest management (Niemelä et al., 2007).